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Titanophoneus potens skull    



Skull of Titanophoneus potens  (after Efremov 1938)

Titanophoneus ("titanic murderer") was a carnivorous dinocephalian therapsid from the Middle to Late Permian period (270-260 mya). The type species,  T. potens, was found at Isheevo in the East European Platform in Russia (Efremov 1938).  They belong to the subphyllum  Synapsida, the order Therapsida, the suborder Dinocephalia, and the family Anteosauridae.  

Dinocephalian ("terrible head") fossils are widespread in Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania. Dinocephalians were large-bodied early therapsids that included herbivores, carnivores, and omnivores, some semi-aquatic and others terrestial. They were among the largest animals of the Permian period 

Titanophoneus had an adult body length of about 2.85 m and a skull length of 80 cm, with a heavy long snout. The long tail and short limbs, representing early therapsid features, show the species to be a primitive therapsid with a sprawling stance. The teeth are large, with twelve large palate incisors followed by two projecting canines and various smaller back teeth. The form of Titanophoneus retains similarities with those of earlier pelycosaur synapsids, such as Dimetrodon.


References

Carroll, R. L. 1988. Vertebrate Paleontology and Evolution. WH Freeman & Co.

Efremov, I.  1938.  Some new Permian reptiles of the USSR, Comptes Rendus Acad. Sci. USSR. Paleontol., Vol 19, N 9.


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