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Charassognathus gracilis skull



Skull of Charassognathus gracilis with labelled bones (after Botha et al. 2007.)


Charassognathus gracilis is the earliest known cynodont ("dog-teeth"), a  therapsid sub-order occuring widely during the Late Permian in South Africa and elsewhere. The name Charassognathus means Charasso- "notched,"  -gnathus "jaw", while gracilis means "slender" or gracile.   It is in the class Synapsida, the order Therapsida, and the family Cynodontia.  C. gracilis was discovered by Botha et al. (2007) near Fraserburg, South Africa in a Late Permian formation dated at 260 mya.      

Charassognathus gracilis was a small herbivore with a skull length of 5 cm (2 in). It is named for a notch at the back of its lower jaw which was used to attach the adductor mandibula externus muscle, used for chewing. Its teeth were diversified, with four types found in cynodonts, including incisors, very large canines, tricuspid premolars (see figure inset "T"), and molars.  Its skull has a large synapsid opening in the temporal region for attachment of the adductor mandibula externus.            
           

References:

Botha et al. 2007  

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